*,
*::before,
*::after {
  margin: 0;
  padding: 0;
  box-sizing: border-box;
}

:root {
  --color-bg: #faf8f4;
  --color-surface: #ffffff;
  --color-text: #2c1810;
  --color-text-secondary: #6b5c52;
  --color-text-light: #9a8b80;
  --color-border: #e8e0d8;
  --color-primary: #2d6b3f;
  --color-primary-light: #e8f5e9;
  --color-accent: #c4912e;
  --color-accent-light: #fef3e0;
  --color-protected: #c0392b;
  --color-protected-light: #fdecea;
  --color-rare: #7b1fa2;
  --color-rare-light: #f3e5f5;
  --color-blue: #1565c0;
  --color-blue-light: #e3f2fd;
  --font-display: 'Playfair Display', Georgia, serif;
  --font-body: 'Inter', -apple-system, BlinkMacSystemFont, 'Segoe UI', sans-serif;
  --shadow-sm: 0 1px 3px rgba(44,24,16,0.06);
  --shadow-md: 0 4px 12px rgba(44,24,16,0.08);
  --shadow-lg: 0 8px 30px rgba(44,24,16,0.12);
  --radius-sm: 6px;
  --radius-md: 10px;
  --radius-lg: 16px;
  --transition: 0.2s ease;
}

html {
  scroll-behavior: smooth;
  scroll-padding-top: 140px;
}

body {
  font-family: var(--font-body);
  background: var(--color-bg);
  color: var(--color-text);
  line-height: 1.6;
  -webkit-font-smoothing: antialiased;
}

/* ============ HERO ============ */

.hero {
  position: relative;
  min-height: 85vh;
  display: flex;
  align-items: center;
  justify-content: center;
  text-align: center;
  background:
    linear-gradient(135deg, #1a3a2a 0%, #2d5016 30%, #4a7c28 60%, #6b9b37 100%);
  overflow: hidden;
}

.hero::before {
  content: '';
  position: absolute;
  inset: 0;
  background:
    radial-gradient(ellipse at 20% 50%, rgba(196,145,46,0.15) 0%, transparent 60%),
    radial-gradient(ellipse at 80% 30%, rgba(255,255,255,0.05) 0%, transparent 50%);
}

.hero-overlay {
  position: absolute;
  inset: 0;
  background: url("data:image/svg+xml,%3Csvg width='60' height='60' viewBox='0 0 60 60' xmlns='http://www.w3.org/2000/svg'%3E%3Cg fill='none' fill-rule='evenodd'%3E%3Cg fill='%23ffffff' fill-opacity='0.03'%3E%3Cpath d='M36 34v-4h-2v4h-4v2h4v4h2v-4h4v-2h-4zm0-30V0h-2v4h-4v2h4v4h2V6h4V4h-4zM6 34v-4H4v4H0v2h4v4h2v-4h4v-2H6zM6 4V0H4v4H0v2h4v4h2V6h4V4H6z'/%3E%3C/g%3E%3C/g%3E%3C/svg%3E") repeat;
}

.hero-content {
  position: relative;
  z-index: 1;
  padding: 2rem;
}

.hero-super {
  font-family: var(--font-body);
  font-size: 0.85rem;
  font-weight: 500;
  letter-spacing: 0.15em;
  text-transform: uppercase;
  color: rgba(255,255,255,0.7);
  margin-bottom: 1rem;
}

.hero h1 {
  font-family: var(--font-display);
  font-size: clamp(2.5rem, 7vw, 5rem);
  font-weight: 700;
  color: #fff;
  line-height: 1.1;
  margin-bottom: 1rem;
  text-shadow: 0 2px 20px rgba(0,0,0,0.2);
}

.hero-subtitle {
  font-family: var(--font-display);
  font-size: clamp(1.1rem, 2.5vw, 1.5rem);
  font-style: italic;
  color: rgba(255,255,255,0.85);
  margin-bottom: 1.5rem;
}

.hero-count {
  display: inline-block;
  font-size: 0.9rem;
  font-weight: 500;
  color: var(--color-accent);
  background: rgba(255,255,255,0.12);
  backdrop-filter: blur(8px);
  padding: 0.5rem 1.5rem;
  border-radius: 100px;
  border: 1px solid rgba(255,255,255,0.15);
}

.hero-scroll-hint {
  position: absolute;
  bottom: 2rem;
  left: 50%;
  transform: translateX(-50%);
  color: rgba(255,255,255,0.5);
  animation: bob 2s ease-in-out infinite;
}

@keyframes bob {
  0%, 100% { transform: translateX(-50%) translateY(0); }
  50% { transform: translateX(-50%) translateY(8px); }
}

/* ============ STICKY NAV ============ */

.sticky-nav {
  position: sticky;
  top: 0;
  z-index: 100;
  background: rgba(250,248,244,0.92);
  backdrop-filter: blur(12px);
  border-bottom: 1px solid var(--color-border);
  transition: box-shadow var(--transition);
}

.sticky-nav.scrolled {
  box-shadow: var(--shadow-md);
}

.nav-inner {
  max-width: 1400px;
  margin: 0 auto;
  padding: 0.75rem 1.5rem;
  display: flex;
  align-items: center;
  gap: 1.5rem;
  flex-wrap: wrap;
}

.nav-brand {
  font-family: var(--font-display);
  font-weight: 600;
  font-size: 1.1rem;
  color: var(--color-primary);
  white-space: nowrap;
}

.nav-controls {
  display: flex;
  align-items: center;
  gap: 1rem;
  flex: 1;
  flex-wrap: wrap;
}

.search-wrap {
  position: relative;
  flex: 1;
  min-width: 200px;
  max-width: 400px;
}

.search-icon {
  position: absolute;
  left: 12px;
  top: 50%;
  transform: translateY(-50%);
  color: var(--color-text-light);
  pointer-events: none;
}

.search-wrap input {
  width: 100%;
  padding: 0.55rem 2.5rem 0.55rem 2.5rem;
  border: 1.5px solid var(--color-border);
  border-radius: 100px;
  font-family: var(--font-body);
  font-size: 0.9rem;
  background: var(--color-surface);
  color: var(--color-text);
  outline: none;
  transition: border-color var(--transition), box-shadow var(--transition);
}

.search-wrap input:focus {
  border-color: var(--color-primary);
  box-shadow: 0 0 0 3px rgba(45,107,63,0.12);
}

.search-wrap input::placeholder {
  color: var(--color-text-light);
}

.search-kbd {
  position: absolute;
  right: 10px;
  top: 50%;
  transform: translateY(-50%);
  font-size: 0.7rem;
  padding: 2px 6px;
  border: 1px solid var(--color-border);
  border-radius: 4px;
  color: var(--color-text-light);
  background: var(--color-bg);
  pointer-events: none;
}

.filter-group {
  display: flex;
  gap: 0.4rem;
}

.filter-btn {
  padding: 0.4rem 0.9rem;
  border: 1.5px solid var(--color-border);
  border-radius: 100px;
  background: transparent;
  font-family: var(--font-body);
  font-size: 0.8rem;
  font-weight: 500;
  color: var(--color-text-secondary);
  cursor: pointer;
  transition: all var(--transition);
  white-space: nowrap;
}

.filter-btn:hover {
  border-color: var(--color-primary);
  color: var(--color-primary);
}

.filter-btn.active {
  background: var(--color-primary);
  border-color: var(--color-primary);
  color: #fff;
}

/* ============ CITY SELECTOR ============ */

.location-wrap {
  position: relative;
  display: flex;
  align-items: center;
}

.location-icon {
  position: absolute;
  left: 10px;
  color: var(--color-primary);
  pointer-events: none;
  z-index: 1;
}

.location-wrap select {
  appearance: none;
  padding: 0.45rem 2rem 0.45rem 2rem;
  border: 1.5px solid var(--color-primary);
  border-radius: 100px;
  font-family: var(--font-body);
  font-size: 0.8rem;
  font-weight: 500;
  color: var(--color-primary);
  background: var(--color-primary-light);
  cursor: pointer;
  outline: none;
  transition: all var(--transition);
  background-image: url("data:image/svg+xml,%3Csvg width='10' height='6' viewBox='0 0 10 6' xmlns='http://www.w3.org/2000/svg'%3E%3Cpath d='M1 1l4 4 4-4' fill='none' stroke='%232d6b3f' stroke-width='1.5' stroke-linecap='round'/%3E%3C/svg%3E");
  background-repeat: no-repeat;
  background-position: right 10px center;
}

.location-wrap select:hover {
  background-color: #d4ecd9;
}

.location-wrap select:focus {
  box-shadow: 0 0 0 3px rgba(45,107,63,0.15);
}

/* ============ LIKELIHOOD BADGES ============ */

.likelihood-badge {
  position: absolute;
  top: 0.75rem;
  right: 0.75rem;
  font-size: 0.6rem;
  font-weight: 600;
  letter-spacing: 0.04em;
  text-transform: uppercase;
  padding: 0.2rem 0.55rem;
  border-radius: 100px;
  z-index: 2;
  backdrop-filter: blur(6px);
}

.likelihood-badge.common {
  background: rgba(45,107,63,0.9);
  color: #fff;
}

.likelihood-badge.occasional {
  background: rgba(196,145,46,0.9);
  color: #fff;
}

.city-habitat-note {
  display: inline-flex;
  align-items: center;
  gap: 0.4rem;
  font-size: 0.82rem;
  color: var(--color-text-secondary);
  background: var(--color-primary-light);
  border: 1px solid rgba(45,107,63,0.15);
  padding: 0.4rem 0.9rem;
  border-radius: var(--radius-sm);
  margin-bottom: 1.5rem;
}

.city-habitat-note svg {
  width: 15px;
  height: 15px;
  color: var(--color-primary);
  flex-shrink: 0;
}

.city-stats {
  font-weight: 600;
  color: var(--color-primary);
}

/* ============ FAMILY NAV ============ */

.family-nav {
  position: sticky;
  top: 60px;
  z-index: 99;
  background: rgba(250,248,244,0.88);
  backdrop-filter: blur(8px);
  border-bottom: 1px solid var(--color-border);
  overflow-x: auto;
  -webkit-overflow-scrolling: touch;
  scrollbar-width: none;
}

.family-nav::-webkit-scrollbar {
  display: none;
}

.family-nav-inner {
  display: flex;
  gap: 0.25rem;
  padding: 0.5rem 1.5rem;
  max-width: 1400px;
  margin: 0 auto;
}

.family-link {
  padding: 0.35rem 0.8rem;
  border-radius: 100px;
  font-size: 0.78rem;
  font-weight: 500;
  color: var(--color-text-secondary);
  text-decoration: none;
  white-space: nowrap;
  transition: all var(--transition);
}

.family-link:hover {
  background: var(--color-primary-light);
  color: var(--color-primary);
}

.family-link.active {
  background: var(--color-primary);
  color: #fff;
}

/* ============ MAIN CONTENT ============ */

main {
  max-width: 1400px;
  margin: 0 auto;
  padding: 2rem 1.5rem 4rem;
}

/* ============ FAMILY SECTIONS ============ */

.family-section {
  margin-bottom: 4rem;
}

.family-header {
  display: flex;
  align-items: baseline;
  gap: 1rem;
  margin-bottom: 0.5rem;
  padding-top: 1rem;
  flex-wrap: wrap;
}

.family-header h2 {
  font-family: var(--font-display);
  font-size: 2rem;
  font-weight: 700;
  color: var(--color-text);
}

.family-header .family-common {
  font-family: var(--font-display);
  font-size: 1.2rem;
  font-style: italic;
  color: var(--color-text-secondary);
}

.family-description {
  font-size: 0.95rem;
  color: var(--color-text-secondary);
  max-width: 800px;
  margin-bottom: 2rem;
  line-height: 1.7;
}

/* ============ SUBFAMILY ============ */

.subfamily-section {
  margin-bottom: 3rem;
}

.subfamily-header {
  display: flex;
  align-items: center;
  gap: 0.75rem;
  margin-bottom: 1.5rem;
  padding-bottom: 0.5rem;
  border-bottom: 2px solid var(--color-border);
}

.subfamily-header h3 {
  font-family: var(--font-display);
  font-size: 1.3rem;
  font-weight: 600;
  color: var(--color-text);
}

.subfamily-header .subfamily-common {
  font-size: 0.9rem;
  color: var(--color-text-light);
}

.subfamily-header .species-count-badge {
  margin-left: auto;
  font-size: 0.75rem;
  font-weight: 600;
  color: var(--color-text-light);
  background: var(--color-bg);
  border: 1px solid var(--color-border);
  padding: 0.2rem 0.6rem;
  border-radius: 100px;
}

/* ============ SPECIES GRID ============ */

.species-grid {
  display: grid;
  grid-template-columns: repeat(auto-fill, minmax(340px, 1fr));
  gap: 1.25rem;
}

/* ============ SPECIES CARD ============ */

.species-card {
  background: var(--color-surface);
  border-radius: var(--radius-lg);
  border: 1px solid var(--color-border);
  overflow: hidden;
  transition: transform var(--transition), box-shadow var(--transition);
  position: relative;
}

.species-card:hover {
  transform: translateY(-3px);
  box-shadow: var(--shadow-lg);
}

.species-card.hidden {
  display: none;
}

.card-image-wrap {
  position: relative;
  width: 100%;
  aspect-ratio: 4 / 3;
  min-height: 180px;
  background: #e8e0d8;
  overflow: hidden;
}

.card-image-wrap img {
  width: 100%;
  height: 100%;
  object-fit: cover;
  transition: transform 0.4s ease, opacity 0.4s ease;
  opacity: 0;
}

.card-image-wrap img.loaded {
  opacity: 1;
}

.species-card:hover .card-image-wrap img.loaded {
  transform: scale(1.05);
}

.image-placeholder {
  position: absolute;
  inset: 0;
  display: flex;
  flex-direction: column;
  align-items: center;
  justify-content: center;
  color: var(--color-text-light);
  gap: 0.5rem;
}

.image-placeholder.loading {
  background: linear-gradient(135deg, #e8e0d8 0%, #d4ccc4 100%);
}

.image-placeholder.loading::after {
  content: '';
  position: absolute;
  inset: 0;
  background: linear-gradient(
    90deg,
    transparent 0%,
    rgba(255,255,255,0.45) 40%,
    rgba(255,255,255,0.45) 60%,
    transparent 100%
  );
  animation: shimmer 1.6s ease-in-out infinite;
}

@keyframes shimmer {
  0%   { transform: translateX(-100%); }
  100% { transform: translateX(100%); }
}

.image-placeholder.loading svg {
  opacity: 0.2;
  animation: pulse 2s ease-in-out infinite;
}

@keyframes pulse {
  0%, 100% { opacity: 0.15; transform: scale(1); }
  50%      { opacity: 0.3;  transform: scale(1.05); }
}

.image-placeholder.loading .loading-label {
  font-size: 0.7rem;
  font-weight: 500;
  letter-spacing: 0.08em;
  text-transform: uppercase;
  color: var(--color-text-light);
  opacity: 0.6;
  background: rgba(255,255,255,0.5);
  padding: 0.2rem 0.7rem;
  border-radius: 100px;
}

.image-placeholder:not(.loading) svg {
  opacity: 0.25;
}

.image-placeholder:not(.loading) .loading-label {
  font-size: 0.75rem;
  opacity: 0.5;
}

.image-credit {
  position: absolute;
  bottom: 0;
  right: 0;
  font-size: 0.65rem;
  padding: 2px 8px;
  background: rgba(0,0,0,0.55);
  color: rgba(255,255,255,0.8);
  border-radius: var(--radius-sm) 0 0 0;
}

.image-credit a {
  color: rgba(255,255,255,0.9);
  text-decoration: none;
}

.image-credit a:hover {
  text-decoration: underline;
}

.card-badges {
  position: absolute;
  top: 0.75rem;
  left: 0.75rem;
  display: flex;
  gap: 0.35rem;
  z-index: 2;
}

.badge {
  font-size: 0.65rem;
  font-weight: 600;
  letter-spacing: 0.04em;
  text-transform: uppercase;
  padding: 0.2rem 0.6rem;
  border-radius: 100px;
  backdrop-filter: blur(6px);
}

.badge-protected {
  background: rgba(192,57,43,0.9);
  color: #fff;
}

.badge-rare {
  background: rgba(123,31,162,0.85);
  color: #fff;
}

.card-body {
  padding: 1.25rem;
}

.card-body h4 {
  font-family: var(--font-display);
  font-size: 1.15rem;
  font-weight: 600;
  color: var(--color-text);
  margin-bottom: 0.15rem;
  line-height: 1.3;
}

.scientific-name {
  font-style: italic;
  font-size: 0.85rem;
  color: var(--color-primary);
  margin-bottom: 0.75rem;
}

.card-description {
  font-size: 0.88rem;
  line-height: 1.65;
  color: var(--color-text-secondary);
  margin-bottom: 1rem;
}

.card-meta {
  display: flex;
  flex-wrap: wrap;
  gap: 0.5rem;
}

.meta-tag {
  display: inline-flex;
  align-items: center;
  gap: 0.3rem;
  font-size: 0.75rem;
  color: var(--color-text-light);
  background: var(--color-bg);
  padding: 0.25rem 0.6rem;
  border-radius: 100px;
}

.meta-tag svg {
  width: 13px;
  height: 13px;
  flex-shrink: 0;
}

/* ============ NO RESULTS ============ */

.no-results {
  text-align: center;
  padding: 4rem 2rem;
  color: var(--color-text-light);
}

.no-results h3 {
  font-family: var(--font-display);
  font-size: 1.5rem;
  margin-bottom: 0.5rem;
  color: var(--color-text-secondary);
}

.no-results p {
  font-size: 0.95rem;
}

/* ============ BACK TO TOP ============ */

.back-to-top {
  position: fixed;
  bottom: 2rem;
  right: 2rem;
  width: 44px;
  height: 44px;
  border-radius: 50%;
  background: var(--color-primary);
  color: #fff;
  border: none;
  cursor: pointer;
  display: flex;
  align-items: center;
  justify-content: center;
  box-shadow: var(--shadow-md);
  opacity: 0;
  visibility: hidden;
  transform: translateY(10px);
  transition: all var(--transition);
  z-index: 50;
}

.back-to-top.visible {
  opacity: 1;
  visibility: visible;
  transform: translateY(0);
}

.back-to-top:hover {
  background: #245532;
  box-shadow: var(--shadow-lg);
}

/* ============ FOOTER ============ */

.site-footer {
  background: var(--color-text);
  color: rgba(255,255,255,0.75);
  padding: 3rem 1.5rem;
}

.footer-inner {
  max-width: 1400px;
  margin: 0 auto;
  display: grid;
  grid-template-columns: 1fr 1fr;
  gap: 3rem;
}

.footer-source h3 {
  font-family: var(--font-display);
  font-size: 1.2rem;
  color: #fff;
  margin-bottom: 0.75rem;
}

.footer-source p {
  font-size: 0.9rem;
  line-height: 1.7;
  margin-bottom: 0.75rem;
}

.footer-source a {
  color: var(--color-accent);
  text-decoration: none;
}

.footer-source a:hover {
  text-decoration: underline;
}

.footer-note {
  font-size: 0.82rem !important;
  opacity: 0.75;
}

.footer-meta {
  display: flex;
  flex-direction: column;
  justify-content: flex-end;
  font-size: 0.85rem;
  line-height: 1.7;
}

.footer-meta a {
  color: var(--color-accent);
  text-decoration: none;
}

.footer-meta a:hover {
  text-decoration: underline;
}

.footer-copy {
  margin-top: auto;
  opacity: 0.6;
  font-size: 0.8rem;
}

/* ============ RESPONSIVE ============ */

@media (max-width: 900px) {
  .footer-inner {
    grid-template-columns: 1fr;
    gap: 2rem;
  }
}

@media (max-width: 768px) {
  html {
    scroll-padding-top: 180px;
  }

  .hero {
    min-height: 70vh;
  }

  .nav-inner {
    padding: 0.6rem 1rem;
  }

  .nav-brand {
    display: none;
  }

  .nav-controls {
    width: 100%;
  }

  .search-wrap {
    max-width: 100%;
  }

  .search-kbd {
    display: none;
  }

  .family-nav {
    top: 52px;
  }

  .family-nav-inner {
    padding: 0.4rem 1rem;
  }

  .species-grid {
    grid-template-columns: 1fr;
  }

  main {
    padding: 1.5rem 1rem 3rem;
  }

  .family-header h2 {
    font-size: 1.5rem;
  }

  .subfamily-header h3 {
    font-size: 1.1rem;
  }

  .back-to-top {
    bottom: 1rem;
    right: 1rem;
  }
}

@media (max-width: 480px) {
  .filter-group {
    width: 100%;
    justify-content: stretch;
  }

  .filter-btn {
    flex: 1;
    text-align: center;
  }
}

/* ============ PRINT ============ */

@media print {
  .sticky-nav,
  .family-nav,
  .back-to-top,
  .hero-scroll-hint {
    display: none !important;
  }

  .hero {
    min-height: auto;
    padding: 2rem;
  }

  .species-card {
    break-inside: avoid;
    border: 1px solid #ccc;
  }
}